Dear Dirk,

Here it is.

All the best,


Dr Jean-Bernard HUCHET, PhD
UMR 7209 du CNRS, Archéozoologie, Archéobotanique
Sociétés, Pratiques et Environnements
Muséum National d'Histoire Naturelle
CP 56, 55 rue Buffon 75005 Paris
33 (0) 1 40 79 32 74
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UMR 7205, Origine, Structure et Evolution de la Biodiversité (OSEB).
Département Systématique et Evolution (Entomologie)
Muséum National d'Histoire Naturelle
45 rue Buffon 75005 Paris
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Laboratoire d'Anthropologie (A3P)
PACEA - UMR 5199, Université Bordeaux 1
Bât. B 8, Avenue des Facultés
33405 Talence Cedex France
Tel : 33 (0) 5 40 00 25 48
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Message du : 15/06/2013 15:24
De : "Dirk " <[log in to unmask]>
A : [log in to unmask]
Copie à :
Sujet : Re: Spangler 1988

Hi All,

Does anyone have a pdf of this?

Antonio Arillo* & Vicente M. Ortuño (2008)  Did dinosaurs have any relation with dung-beetles? (The origin of coprophagy). Journal of Natural History 42, Issue 19-20: p. 1405-1408.

Thanks in advance,


On 14 Jun 2013, at 17:22, Cole Gilbert wrote:

HI Clarke,
There are also scarabs that have a tympanic ear, as described by my colleagues in the attached pdf. It includes the reference to Paul Spangler’s paper on cicindelids.
Cole Gilbert
From: Scarab Beetle Taxonomy Discussion List [mailto:[log in to unmask]] On Behalf Of Clarke Scholtz
Sent: Friday, June 14, 2013 11:09 AM
To: [log in to unmask]
Subject: Spangler 1988
Dear Colleagues,
I am looking for a reference by Paul Spangler in which he stated that tiger beetles are one of few beetle groups with a hearing tympanum.
I have no idea what the title of the paper was, or where it was published, but if anyone can help me locate it (or send me a copy!), I’d be most grateful.
Best wishes,
Professor Clarke Scholtz
Scarab Research Group
Department of Zoology & Entomology
University of Pretoria
Pretoria 0002 - RSA
Fax +27 12 362 5242
Phone +27 12 420 3236
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<Scarab tympanum Forrest et al 1997..pdf>