I'm not sure that (m)any Staphyliniformia have such deep and complex sculpture on the elytra? If I'm right about squamosity, then it pretty much rules out Staphyliniformia also ...

From: Max Barclay <[log in to unmask]>
To: [log in to unmask]
Sent: Sunday, 23 June 2013 12:50 AM
Subject: Re: Pleistocene scarab fragments?

Look at the apex of the elytra.. that looks like an elytron that doesn't cover the whole abdomen, i.e. leaves some segments visible from above. I have no idea how big they are of course, but I would have started with Staphyliniformia- some histeroid or omaliine or something.. that doesn't say scarab or weevil to me..


Maxwell V. L. Barclay
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From: Scarab Beetle Taxonomy Discussion List [[log in to unmask]] on behalf of Paul Schoolmeesters [[log in to unmask]]
Sent: 22 June 2013 13:34
To: [log in to unmask]
Subject: Re: Pleistocene scarab fragments?

It looks a bit like Euparixoides, but these are South American species

Paul Schoolmeesters
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Coprophagous Scarabaeidae of the world

From: Scarab Beetle Taxonomy Discussion List [mailto:[log in to unmask]] On Behalf Of Frank T. Krell
Sent: samedi 22 juin 2013 00:40
To: [log in to unmask]
Subject: Pleistocene scarab fragments?

Hi all,

I am a little bit lost, so it’s time to ask the list, I guess.
I am working on scarab fragments of a Pleistocene site of Colorado. Most of them are Aphodius heads (or Aphodius s.l. heads, for the splitters). I also got a few elytral fragments with a pretty distinct sculpture, consisting of broad deep, largely punctured stripes and not much broader, elevated, equally largely punctured intervals (see attached photo).
I thought I have seen such elytra before, but cannot remember. I got them as possible scarabs, but I am struggling to find a North American scarab with such elytra. Could it be something else? I cc to Mike Ivie because he knows all those little families that nobody else knows (or are you on the Scarbs-L list, Mike?).
Any ideas? Thanks!


Dr. Frank-T. Krell
Curator of Entomology
Commissioner, International Commission on Zoological Nomenclature
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Department of Zoology
Denver Museum of Nature & Science
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