Others are:
Polyphylla (s. str.) fullo (Linnaeus, 1758)
Miltotrogus vernus Germar, 1823.
Source: http://elibrary.lt/resursai/Uzsienio%20leidiniai/MFTI/2003/041.pdf
Codocera ferruginea Eschscholtz, 1818
Citation: Bezborodov V.G. Annotated list of the lamellicorn beetles (Coleoptera, Scarabaeoidea) of the fauna of Primorsky Krai (Russia). Amurian zoological journal. VI(1), 2014: 22-50
Суббота, 14 марта 2020, 20:54 +03:00 от Dirk Ahrens <[log in to unmask]>:

Hi All,

in a work in preparation we would like to give in a few sentences a short review on non-feeding scarab adults.
Could you guys give us a help to make this thing more complete?
There are in literature some vague statements such as this of Ritcher (1958) and Scholtz (1990) about nonfeeding Rutelinae and Dynastinae, for which I could find so far no precise data on taxa and references. Maybe I have overlooked other references…
In order to avoid notification already known to us, I attached below the list I could make so far…
Thanks to all in advance who may help!
ps. Do we have this pattern (i.e., nonfeeding of adult specimens) also among dung beetle lineages (Aphodiinae/ Scarabaeinae)?


Non-feeding in Scarabaeoidea


·      Diphyllostoma (Lucanidae; nonfunctional mandibles; maxillae with a small rounded galea; Nel & Scholtz 1990)

·      Amphicoma, Lichnanthe (Glaphyridae; Ritcher 1958; Sabatinelli et al. 2020)

·      Pleocoma (Pleocomidae; Ritcher 1958; nonfunctional mandibles; maxillae with a small rounded galea; Nel & Scholtz 1990

·      Pachypus (Pachypodiini; Crovetti 1968)

·      Ceramida (Tanyproctini; very simple and reduced mouthparts; Sanmartin & Martin-Piera 2000; some more candidates with reduced epipharynx: Alaia, Atanyproctus, and Otoclinius)

·      Aclopus (Aclopinae; reduced galea, simple epipharynx; Nel & Scholtz 1990)

·      Xenaclopus (Melolonthinae, s.l. (Ocampo & Vaz de Mello 2008): reduced galea, simple epipharynx; Nel & Scholtz 1990)

·      Scapanoclypeus ("Tanyproctini" (see Eberle et al. 2018); epipharynx reduced, galea, lacinia, ligula absent; Nel & Scholtz 1990)

·      Sebaris, Pseudachloa, Eucyclophylla (Melolonthinae, s.l.; galea oval with setae; group 4 and 5; Nel & Scholtz 1990)

·      "There are species in each of the subfamilies Melolonthinae, Rutelinae, and Dynastinae which do not feed at all in the adult stage" (without specifications; Ritcher 1958, Scholtz 1990)

All the best, Dr. Mikhail F. Bagaturov

IUCN/SSC Amphibian SG and Conservation Planning SG, IPBES 
Zoological Institute RAS, St. Petersburg, Russia/ Insect Taxo Lab.,
Academic Member of Athens Institute For Education & Research (http://atiner.gr)

Author of Theraphosids of the World: http://tarantulas.su/